We present U-Pb zircon age determinations of two Variscan ultrapotassic plutonic rocks from the Moldanubian Zone (Bohemian Massif). Equant, multifaceted zircons without inherited cores from a two-pyroxene-biotite quartz monzonite of the Jihlava Pluton yielded a precise age of 335.12 +/- 0.57 Ma, interpreted as dating magma crystallization. The majority of both tabular and prismatic grains from the amphibole-biotite melagranite ("durbachite") from the Trebic Pluton plot along a discordia intersecting the concordia at 334.8 +/- 3.2 Ma; prismatic zircon grains commonly contain inherited cores and yield an upper intercept age of 2.2 Ga, indicating early Proterozoic inheritance. We therefore suggest that both types of the ultrapotassic plutonic rocks from the Bohemian Massif crystallized at ca 335 Ma, and the previously published ages higher than ca 340 Ma for "durbachites" were biased by a small amount of unresolved inheritance. The ultrapotassic magma emplacement in the middle crust was related to rapid exhumation of a deep crustal segment, considered as isothermal decompression between high-pressure (similar to 340 Ma) and medium-pressure (similar to 333 Ma) stages recorded in granulites. Mineral assemblages as well as external and internal zircon morphology suggest that the Jihlava intrusion was deep and dry, whereas the Trebic intrusion was shallow and wet. Low epsilon(Hf) values of zircons (-4.4 to -7.5) in both rock types suggest a similar source with a predominant crustal component. However, inherited grains in the Trebic melagranite indicate its contamination with crustal material during emplacement, and thus possibly a slower rate of exhumation and/or of magma ascent through the crust